Protein Modification Ontology (MOD)

Name: Protein Modification Ontology (MOD)
Published: 2026-11-13T13:50:45.374984Z

Selected English labels and exact synonyms from the Protein Modification Ontology (MOD), retrieved 2026-11-07, with source identifiers and spelling preserved. Licensed CC BY 4.0; full attribution is listed under Sources. Under semantic review; not endorsed by the ontology's maintainers.

Curator: Oliver Bennett

Languages shown reflect this page; source translations may cover only some concepts.

English
6-[S-cysteinyl]flavin mononucleotideUnrated 6-[S-cysteinyl]FMNUnrated MOD_RES S-6-FMN cysteineUnrated (R)-2-amino-3-[6-riboflavin 5'-dihydrogen phosphate]sulfanylpropanoic acidUnrated S6FMNCysUnrated S-(6-FMN)-L-cysteineUnrated
hydrolyzed N-methylmaleimide cysteine adductUnrated
2-amino-3-methoxyprop-2-enoic acidUnrated 3-methoxydehydroalanineUnrated 3-methoxydidehydroalanineUnrated
(2S)-2-amino-3-(6-bromo-1H-indol-3-yl)propanoic acidUnrated 6'-bromo-L-tryptophanUnrated 6'-BrTrpUnrated MOD_RES 6'-bromotryptophanUnrated
(2S)-2-amino-3-(2-acetamido-2-deoxy-alpha-D-galactopyranosyloxy)propanoic acidUnrated CARBOHYD O-linked (GalNAc) serineUnrated mucin type O-glycosylserineUnrated O3-(N-acetylgalactosaminyl)serineUnrated OGalNAcSerUnrated O-(N-acetylamino)galactosyl-L-serineUnrated
KResUnrated potassium containing modified residueUnrated
2-hydroxyglycineUnrated 2-hydroxyglycine observational artifactUnrated 2HyGlyUnrated alpha-hydroxyglycineUnrated amino(hydroxy)acetic acidUnrated aminohydroxyacetic acidUnrated
D-alanineUnrated D-alanine (Ser)Unrated D-Ala(Ser)Unrated MOD_RES D-alanine (Ser)Unrated (R)-2-aminopropanoic acidUnrated
O-hexosylated threonineUnrated
5-glutaminyl glutamic acidUnrated
monofluorinated L-tyrosineUnrated phefUnrated
Sulfanilic Acid (SA), light C12Unrated
anisyl modified residueUnrated
(2R)-2-amino-3-thiocyanatopropanoic acidUnrated alpha-amino-beta-thiocyanatopropionic acidUnrated beta-thiocyanatoalanineUnrated MOD_RES S-cyanocysteineUnrated S-cyanocysteineUnrated S-cyano-L-cysteineUnrated serine thiocyanic acid esterUnrated
residue reporter fragmentUnrated
t-butyloxymethyl (Bum)Unrated
N2Me3AlaUnrated N,N,N-trimethyl-L-serine (from L-serinium)Unrated
glyglykUnrated N6-glycylglycyl-L-lysineUnrated N6-(glycylglycyl)lysineUnrated ubiquitination signature dipeptidyl lysineUnrated
alpha-amino protonated residueUnrated N2H+NResUnrated
Hex7HexNAc2 N4-glycosylated asparagineUnrated
MOD_RES S-(2,3-dicarboxypropyl)cysteineUnrated S-methylbutanedioic acid-L-cysteineUnrated
(2-aminosuccinimidyl)acetic acidUnrated
3-[(2,6-dimethylpiperidin-1-yl)acetyl]amino)propanoylUnrated TMT6plex-126 reporter+balance reagent O3-acylated threonineUnrated
4,4'-dimethoxybenzhydrylUnrated bis(4-methoxyphenyl)methylUnrated dimethoxybenzhydryl modified residueUnrated MbhUnrated
L-alanine residue (Thr)Unrated Thr(Ala)Unrated
5-oxothiomorpholine-3-carboxylic acidUnrated cyclized N-terminal S-carboxymethyl-L-cysteineUnrated OtcUnrated (R)-5-oxoperhydro-1,4-thiazine-3-carboxylic acidUnrated
residue modified by radical mechanismUnrated
gamma-glutamylpolyglycineUnrated L-isoglutamyl-polyglycineUnrated MOD_RES 5-glutamyl polyglycineUnrated
O-glycosyl-L-threonineUnrated OGlycoThrUnrated
NGlycoResUnrated N-glycosylated residueUnrated
2-aminopentanimideUnrated 3-amino-2,6-piperidinedioneUnrated (3S)-3-aminopiperidine-2,6-dioneUnrated alpha-aminoglutarimideUnrated L-glutamimideUnrated
METAL Iron-sulfur (2Fe-2S)Unrated METAL Iron-sulfur (2Fe-2S); shared with dimeric partnerUnrated tetrakiscysteinato-1kappa(2)S,2kappa(2)S-di-mu-sulfido-diironUnrated tetrakis-L-cysteinyl diiron disulfideUnrated
pyridyl thiol modified L-serineUnrated
reduced 4-hydroxynonenal adductUnrated
modified L-tryptophan residueUnrated ModTrpUnrated
(4-methylpiperazin-1-yl)acetylUnrated iTRAQ8plex-119 reporter+balance reagent O3-acylated serineUnrated
3-[(2,6-dimethylpiperidin-1-yl)acetyl]amino)propanoylUnrated TMT6plex-127 reporter+balance reagent O3-acylated threonineUnrated
(2S)-2-amino-6-([(3S)-3-amino-3-carboxypropanoyl]amino)hexanoic acidUnrated beta-(N6-lysyl)aspartyl acidUnrated CROSSLNK Isoaspartyl lysine isopeptide (Lys-Asn)Unrated isoaspartyl N6-lysineUnrated N6-(L-isoaspartyl)-L-lysineUnrated N6-(L-isoaspartyl)-L-lysine (Asn)Unrated N-(beta-Aspartyl)-Lysine (Crosslink)Unrated N(epsilon)-(beta-aspartyl)lysineUnrated XLNK-4Asp-N6Lys(Asn)Unrated
6-[(1R)-1-amino-2-sulfanylethyl]pyridine-2,5-dicarboxylic acidUnrated L-cysteine pyridine-2,5-dicarboxylic acidUnrated
potassium L-aspartateUnrated
(2S)-2-amino-5-([(2R)-1-([2-([(2R)-2-amino-2-carboxyethyl]sulfanyl)-2-oxoethyl]amino)-1-oxo-3-sulfanylpropan-2-yl]amino)-5-oxopentanoic acidUnrated ACT_SITE S-(gamma-glutamyl-cysteinyl-glycyl)-cysteine intermediateUnrated S-(gamma-glutamyl-cysteinyl-glycyl)-cysteineUnrated S-(glutathion-1-yl)-L-cysteineUnrated
methylamine Michael addition derivatized threonineUnrated
crosslinked D-serine residueUnrated XlnkDSerUnrated
metal or metal cluster coordinated L-glutamine residueUnrated
crosslinked L-valine residueUnrated XlnkValUnrated
(2S)-2-amino-3-(hexadecanoyloxy)propanoic acidUnrated ACT_SITE O-palmitoyl serine intermediateUnrated LIPID O-palmitoyl serineUnrated L-serine hexadecanoate esterUnrated O3-palmitoyl-serineUnrated O-hexadecanoylated L-serineUnrated O-palmitoylated L-serineUnrated O-palmitoyl-L-serineUnrated OPamSerUnrated palmitoylationsUnrated serine palmitate esterUnrated
3-[(2,6-dimethylpiperidin-1-yl)acetyl]amino)propanoylUnrated TMT6plex-129 reporter+balance reagent acylated residueUnrated
mTRAQ reporter fragmentUnrated
HexNAcUnrated N-acetylhexosaminylated residueUnrated
(2S,3S,5Xi)-5-hydroxy-3-methylpyrrolidine-2-carboxylic acidUnrated 5Hy3MeProUnrated 5-hydroxy-3-methyl-L-prolineUnrated 5-hydroxy-3-methylprolineUnrated beta-methyl-delta-hydroxyprolineUnrated MOD_RES 5-hydroxy-3-methylproline (Ile)Unrated
crosslinked L-leucine residueUnrated XlnkLeuUnrated
NOAc2SerUnrated N,O-diacetylated L-serineUnrated N,O-diacetyl-L-serineUnrated N,O-diacetylserineUnrated (S)-2-acetamido-3-acetyloxypropanoic acidUnrated
L-selenocysteinyl tungsten bis(molybdopterin guanine dinucleotide) (Sec)Unrated
1x(13)C,3x(2)H labeled monomethylated L-arginineUnrated
(4-methylpiperazin-1-yl)acetylUnrated iTRAQ8plex-116 reporter+balance reagent O3-acylated serineUnrated
N-ethyl iodoacetamide-d5Unrated
S-(sn-1-glyceryl)-L-cysteineUnrated
phosphorus containing modified residueUnrated
gamma-glutamyl semialdehydeUnrated glutamyl 5-semialdehydeUnrated glutamyl aldehydeUnrated glutamyl semialdehyde (Pro)Unrated Oxidation of proline to gamma-glutamyl semialdehydeUnrated
(13)C labeled residueUnrated
(2S)-2-amino-5-[(2S)-2-amino-2-carboxyethoxy]-5-oxopentanoic acidUnrated CROSSLNK isoglutamyl serine ester (Ser-Glu)Unrated O3-(isoglutamyl)-serineUnrated O(beta)-(gamma-glutamyl)serineUnrated O-gamma-Glutamyl- (Crosslink to Serine)Unrated O-(L-isoglutamyl)-L-serineUnrated O-(L-isoglutamyl)-L-serine (Glu-Ser)Unrated
quaternary amine labeling reagent heavy form (+6amu) (N-term & K)Unrated
Hex2HexNAc2Pent1 N4-glycosylated asparagineUnrated
halogenated phenylalanineUnrated HalPheUnrated
2-carbamic-propanoic acidUnrated N-carboxy-L-alanineUnrated N-carboxymethionineUnrated (S)-2-carboxyamino-propanoic acidUnrated
5-oxothiomorpholine-3-carboxylic acidUnrated cyclized N-terminal S-carboxamidomethyl-L-cysteineUnrated OtcUnrated (R)-5-oxoperhydro-1,4-thiazine-3-carboxylic acidUnrated
2-amino-3-(2-amino-2-carboxyethyl)sulfanylpropanoic acidUnrated (2R,2'R)-3,3'-sulfanediylbis(2-aminopropanoic acid)Unrated 3,3'-thiobis-L-alanineUnrated L-lanthionineUnrated L-lanthionine (Cys-Ser)Unrated (R,R)-2,6-diamino-4-thiaheptanedioic acidUnrated (R,R)-3,3'-thiobis-(2-aminopropanoic acid)Unrated (R,R)-bis(2-amino-2-carboxyethyl)sulfideUnrated (R)-S-(2-amino-2-carboxyethyl)-L-cysteineUnrated XLNK-SCys-(L)3DhaUnrated
natural, standard, encoded residue substitutionUnrated
2-amino-6-(5-[1,2-dithiolan-3-yl]-1-oxopentyl)aminohexanoic acidUnrated (2S)-2-amino-6-(5-[(3R)-1,2-dithiolan-3-yl]pentanamido)hexanoic acidUnrated (2S)-2-amino-6-[(5-[(3R)-1,2-dithiolan-3-yl]pentanoyl)amino]hexanoic acidUnrated (2S,6'R)-2-amino-6-(6,8-dithiooctanamido)hexanoic acidUnrated lipoylkUnrated MOD_RES N6-lipoyllysineUnrated N6-6,8-dithiooctanoyllysineUnrated N6LipLysUnrated N6-lipoyl-L-lysineUnrated N6-lipoyllysineUnrated N-Lipoyl- (on Lysine)Unrated
5-imidazolinone ring crosslinked residues (Gly)Unrated
4-hydroxy-L-tryptophanUnrated (S)-2-Amino-3-(4-hydroxy-1H-indol-3-yl)propanoic acidUnrated
2-acetylamino-3-mercaptopropanoic acidUnrated 2-acetylamino-3-sulfanylpropanoic acidUnrated 2-acetylazanyl-3-sulfanylpropanoic acidUnrated (2R)-2-acetamido-3-sulfanylpropanoic acidUnrated MOD_RES N-acetylcysteineUnrated NAcCysUnrated N-acetylated cysteineUnrated N-acetylcysteineUnrated N-acetyl-L-cysteineUnrated
cysteinato-8kappaS-histidino-1kappaN(tau)-[(2R)-4-carboxy-2-(carboxymethyl)-2-oxidobutanoate-1kappaO(1),1kappaO(2)]-mu6-carbido-2:3:4:5:6:7kappa(6)C-hexa-mu3-sulfido-1:2:3kappa(3)S;1:2:4kappa(3)S;1:3:4kappa(3)S;5:6:8kappa(3)S;5:7:8kappa(3)S;6:7:8kappa(3)S-tri-mu2-sulfido-2:5kappa(2)S;3:6kappa(2)S;4:7kappa(2)S molybdenum heptaironUnrated L-cysteinyl-L-histidino-homocitryl molybdenum heptairon nonasulfideUnrated L-cysteinyl-L-histidino-homocitryl molybdenum heptairon nonasulfide carbideUnrated nitrogenase iron-molybdenum cofactorUnrated
2-amino-3-(sulfothio)propanoic acidUnrated (2R)-2-amino-3-(sulfosulfanyl)propanoic acidUnrated 3-(sulfosulfanyl)-L-alanineUnrated cysteine-S-sulfonic acidUnrated cysteine sulfate thioesterUnrated SSulfCysUnrated S-sulfocysteineUnrated S-sulfo-L-cysteineUnrated
glutamyl semialdehydeUnrated L-glutamic gamma-semialdehydeUnrated (S)-2-amino-5-oxopentanoic acidUnrated
OFoResUnrated O-formylated residueUnrated
3-[(2,6-dimethylpiperidin-1-yl)acetyl]amino)propanoylUnrated TMT6plex-131 reporter+balance reagent O3-acylated threonineUnrated
Michael addition of BHT quinone methide to cysteine and lysineUnrated
(4-methylpiperazin-1-yl)acetylUnrated iTRAQ8plex-115 reporter+balance reagent O3-acylated threonineUnrated
2-amino-3-[(S)-2-((Z)-9-octadecenoyloxy)-3-(hexadecanoyloxy)propyl]thiopropanoic acidUnrated (2R)-2-amino-3-[(2S)-2-((9Z)-9-octadecenoyloxy)-3-(hexadecanoyloxy)propyl]sulfanylpropanoic acidUnrated LIPID S-diacylglycerol cysteineUnrated S-(1-2'-oleoyl-3'-palmitoyl-glycerol)cysteineUnrated S-(2',3'-diacylglycerol)-L-cysteineUnrated SAcyl2GlyceroCysUnrated S-diacylglycerol-L-cysteineUnrated
(2S)-2-amino-3-(5,6-dihydroxy-1H-indol-3-yl)propanoic acidUnrated 5,6-DihydroxytryptophanUnrated Double oxidation of TrpUnrated
S-(2-mercaptoethyl)cysteineUnrated S-(2-sulfanylethyl)cysteine (Ser)Unrated
di-mu-sulfido(N(eta1)-arginyl-S-cysteinyliron)(bis-S-cysteinyliron)Unrated METAL Iron-sulfur (2Fe-2S)Unrated tris-L-cysteinyl L-arginyl diiron disulfideUnrated
(2S,4R)-4-[2-acetamido-2-deoxy-alpha-D-glucopyranosyloxy]pyrrolidine-2-carboxylic acidUnrated 4-[(2-N-acetylamino)-alpha-D-glucopyranosyl]oxyprolineUnrated 4-(N-acetylglucosaminyloxy)prolineUnrated alpha-2-(N-acetylamino)glucopyranosyl-4-hydroxyprolineUnrated CARBOHYD O-linked (GlcNAc) hydroxyprolineUnrated O4GlcNAcHyProUnrated O4-glycosyl-hydroxyprolineUnrated O4-(N-acetylamino)glucosyl-L-hydroxyprolineUnrated
(2S)-2-[(dihydroxymethyl)amino]-4-(methylsulfanyl)butanoic acidUnrated N-(dihydroxymethyl)-L-methionineUnrated N-(dihydroxymethyl)-L-methionine (Met)Unrated N-formyl-L-methionine hydrateUnrated N-orthoformylmethionineUnrated
N-palmitoyl-S-diacylglycerol-L-cysteineUnrated
(4-methylpiperazin-1-yl)acetylUnrated iTRAQ4plex reporter+balance reagent acylated residue, average mass modificationUnrated
2-(L-cystein-S-yl)-D-tyrosineUnrated (2R,5S)-2,5-diamino-5-carboxyl-6-(hydroxyphenyl)-4-thiahexanoic acidUnrated (2S)-2-amino-2-([(2R)-2-amino-2-carboxyethyl]sulfanyl)-3-(4-hydroxyphenyl)propanoic acidUnrated alpha-(L-cystein-S-yl)-D-tyrosineUnrated
(2S,3S)-2,4-diamino-3-hydroxy-4-oxobutanoic acidUnrated (2S,3S)-2-amino-3-hydroxy-4-butanediamic acidUnrated (2S,3S)-3-hydroxyasparagineUnrated (3S)3HyAsnUnrated L-threo-beta-hydroxyasparagineUnrated MOD_RES (3S)-3-hydroxyasparagineUnrated
(2S)-2-amino-3-(3,4-dihydroxyphenyl)propanoic acidUnrated 3,4-Dihydroxy-Phenylalanine (from Tyrosine) (DOPA)Unrated 3HyTyrUnrated hydroxylationyUnrated L-3',4'-dihydroxyphenylalanineUnrated L-3'-hydroxytyrosineUnrated L-DOPAUnrated levodopaUnrated mod194Unrated MOD_RES 3',4'-dihydroxyphenylalanineUnrated
(2S)-2-amino-3-(4-hydroxyphenyl)propanoic acidUnrated 4-hydroxy-L-phenylalanineUnrated L-tyrosineUnrated L-TyrosineUnrated Phe->TyrUnrated
2-amino-N4,N4-dimethylbutanediamic acidUnrated (2S)-2-amino-4-(dimethylamino)-4-oxobutanoic acidUnrated MOD_RES N4,N4-dimethylasparagineUnrated N4Me2AsnUnrated N4,N4-dimethylated L-asparagineUnrated N4,N4-dimethyl-L-asparagineUnrated N(gamma),N(gamma)-dimethylasparagineUnrated
carbon monoxide dehydrogenase nickel-iron cofactorUnrated hybrid nickel-triiron clusterUnrated mu-1:2kappaO-oxido-mu-1:3kappaO-oxido-mu-2:4kappaO-oxido-mu-3:4kappaS-sulfido-mu3-2:3:4kappaS-sulfido-S-cysteinyl-N1'-histidino-O5-glutamato 1-iron-S5-cysteine persulfido-O3-serinyl 2-nickel-3,4-bis-(S-cysteinyl iron)Unrated Ni-3Fe-2S-3O clusterUnrated tris-L-cysteinyl L-cysteine persulfido L-glutamato L-histidino L-serinyl nickel triiron disulfide trioxideUnrated
O-benzyl-L-threonineUnrated
(79)Br labeled 3',5'-dibromotyrosineUnrated
N6-timnodonoyl-L-lysineUnrated
L-lanthionineUnrated
alpha-amino acidUnrated natural residueUnrated ResUnrated
crosslinked L-cysteine residueUnrated XlnkCysUnrated
2-amino-6-ureido-hexanoic acidUnrated carbamylkUnrated homocitrullineUnrated MOD_RES N6-carbamoyllysineUnrated N6-(aminocarbonyl)-L-lysineUnrated N6-carbamoyl-L-lysineUnrated N6CbmLysUnrated

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