Protein Modification Ontology (MOD)

Name: Protein Modification Ontology (MOD)
Published: 2026-11-13T13:50:45.374984Z

Selected English labels and exact synonyms from the Protein Modification Ontology (MOD), retrieved 2026-11-07, with source identifiers and spelling preserved. Licensed CC BY 4.0; full attribution is listed under Sources. Under semantic review; not endorsed by the ontology's maintainers.

Curator: Oliver Bennett

Languages shown reflect this page; source translations may cover only some concepts.

English
CmCO2Unrated S-carboxymethyl-L-cysteine sulfoneUnrated
ubiquitinylated lysineUnrated
palmitoleylated residueUnrated
MeGluUnrated methylated glutamic acidUnrated
(81)Br labeled 3'-bromotyrosineUnrated
1'-phospho-L-histidineUnrated 2-azanyl-3-(1-phosphono-1H-imidazol-4-yl)propanoic acidUnrated (2S)-2-amino-3-(1-phosphono-1H-imidazol-4-yl)propanoic acidUnrated ACT_SITE Tele-phosphohistidine intermediateUnrated histidine-N1'-phosphateUnrated histidine-N(epsilon)-phosphateUnrated MOD_RES Tele-phosphohistidineUnrated N1-phosphonohistidineUnrated NE2-phosphonohistidineUnrated N(tau)-phosphohistidineUnrated Ntau-phosphorylated L-histidineUnrated NtPHisUnrated tele-phosphohistidineUnrated
METAL Iron-sulfur (4Fe-4S)Unrated METAL Iron-sulfur (4Fe-4S); via tele nitrogenUnrated tetra-mu3-sulfidotris(S-cysteinyliron)(N1'-histidinoiron)Unrated tris-L-cysteinyl L-N1'-histidino tetrairon tetrasulfideUnrated
crosslinked L-methionine residueUnrated XlnkMetUnrated
protonated L-serine (L-serinium) residueUnrated
METAL Iron-sulfur (3Fe-4S)Unrated mu3-sulfido-tri-mu-sulfido-triscysteinato-1kappaS,2kappaS,3kappaS-triironUnrated mu3-sulfido tri-mu-sulfido tris-S-L-cysteinyl triironUnrated tris-L-cysteinyl triiron tetrasulfideUnrated tris-L-cysteinyl triiron tetrasulfide C3 clusterUnrated tris-L-cysteinyl triiron tetrasulfide cubane formUnrated tris-L-cysteinyl triiron tetrasulfide cuboid clusterUnrated tris-L-cysteinyl triiron tetrasulfide trigonal clusterUnrated
diiodinated residueUnrated I2ResUnrated
1'-(8alpha-FAD)-L-histidineUnrated 8alpha-(N1'-histidyl)FADUnrated 8alpha-(N(epsilon)-histidyl)FADUnrated MOD_RES Tele-8alpha-FAD histidineUnrated Nt8aFADHisUnrated N(tau)-(8alpha-FAD)-histidineUnrated N theta and N pi-(8alpha-Flavin) (on Histidine)Unrated (S)-2-amino-3-(1-[8alpha riboflavin 5'-(trihydrogen diphosphate) 5'->5'-ester with adenosine]imidazol-4-yl)propanoic acidUnrated tele-(8alpha-FAD)-histidineUnrated
2x(2)H monomethylated L-lysineUnrated
F1ResUnrated monofluorinated residueUnrated
N-myristoylated residueUnrated NMyrResUnrated
9x(13)C labeled L-tyrosineUnrated
fluorescein-5-thiosemicarbazide - site PUnrated
mod192Unrated NPhosHisUnrated phosphohistidineUnrated phosphorylated L-histidineUnrated
2-amino-butanimideUnrated (3S)-3-amino-2,5-pyrrolidinedioneUnrated alpha-aminosuccinimideUnrated ASIUnrated L-2-aminosuccinimideUnrated L-asparaginimideUnrated L-aspartimideUnrated Succinimide formation from asparagineUnrated
4-aminobutylated residueUnrated
3-[(2,6-dimethylpiperidin-1-yl)acetyl]amino)propanoylUnrated TMT6plex-127 reporter+balance reagent N'-acylated histidineUnrated
[3-[(S)-(4-amino-4-carboxy)butanoyloxymethyl]-7,12-diethenyl-8,13,17-trimethyl-21H,23H-porphine-2,18-bis(2-carboxyethyl)-N21,N22,N23,N24]-ferrateUnrated 5-hydroxymethyl protoporphyrin IX 5-glutamate esterUnrated BINDING Heme (covalent; via 1 link)Unrated cytochrome P450 CYP4A family heme cofactorUnrated hydroxyheme-L-glutamate esterUnrated
quaternary amine labeling reagent heavy form (+6amu) N6-L-lysineUnrated
lysine 4-hydroxynonenal adductUnrated
oxazole/oxazoline ring crosslinked residues (Cys)Unrated
NMeResUnrated N-methylated residueUnrated
3-[(2,6-dimethylpiperidin-1-yl)acetyl]amino)propanoylUnrated TMT6plex-128 reporter+balance reagent O3-acylated serineUnrated
2-acetylamino-4-(methylsulfanyl)butanoic acidUnrated 2-acetylamino-4-(methylthio)butanoic acidUnrated 2-acetylazanyl-4-(methylsulfanyl)butanoic acidUnrated (2S)-2-acetamido-4-(methylsulfanyl)butanoic acidUnrated acetylmethionineUnrated AcMetUnrated methionamineUnrated MOD_RES N-acetylmethionineUnrated N-acetylated L-methionineUnrated N-acetyl-L-methionineUnrated
ribonucleic acid linked residueUnrated RNAResUnrated
Bruker Daltonics SERVA-ICPL(TM) quantification chemistry, heavy form - site KUnrated
2-oxopyrrolidine-5-carboxylic acidUnrated 2-pyrrolidone-5-carboxylic acidUnrated 2-pyrrolidone-5-carboxylic acid (Glu)Unrated (2S)-5-oxo-2-pyrrolidinecarboxylic acidUnrated 5-oxoprolineUnrated 5-oxopyrrolidine-2-carboxylic acidUnrated 5-pyrrolidone-2-carboxylic acidUnrated MOD_RES Pyrrolidone carboxylic acid (Glu)Unrated ntermpeppyroeUnrated PCAUnrated PyrGlu(Glu)Unrated pyroglutamic acidUnrated Pyroglutamic Acid formed from Glutamic AcidUnrated
ntermpeptioacetylUnrated thioacylation of primary amines - site N-termUnrated
hydrolyzed N-ethylmaleimide lysine adductUnrated
6x(13)C labeled L-arginineUnrated arg-13c6Unrated
4-sulfophenyl isothiocyanate N6-derivatized lysineUnrated N6-[(4-sulfophenyl)carbamothioyl]lysineUnrated
(4-methylpiperazin-1-yl)acetylUnrated iTRAQ8plex-115 reporter+balance reagent acylated residueUnrated
quaternary amine labeling reagent light form N6-L-lysineUnrated
fluorescein-5-thiosemicarbazide modified residueUnrated
naphthalene-2,3-dicarboxaldehydeUnrated
hydroxylated isoleucineUnrated HyIleUnrated
2',3'-dihydrotryptophanUnrated
S6FADCysUnrated S-(6-FAD)-L-cysteineUnrated
9-fluorenylmethyloxycarbonyl (Fmoc)Unrated
2-azanyl-3-(3-phosphono-3H-imidazol-4-yl)propanoic acidUnrated (2S)-2-amino-3-(3-phosphono-3H-imidazol-4-yl)propanoic acidUnrated 3'-phospho-L-histidineUnrated ACT_SITE Pros-phosphohistidine intermediateUnrated histidine-N3'-phosphateUnrated histidine-N(delta)-phosphateUnrated MOD_RES Pros-phosphohistidineUnrated N3-phosphonohistidineUnrated ND1-phosphonohistidineUnrated N(pi)-phosphohistidineUnrated Npi-phosphorylated L-histidineUnrated NpPHisUnrated pros-phosphohistidineUnrated
polysulfide crosslinked residuesUnrated
2,5-dimethylpyrrole lysine from 2,5-hexanedione adductUnrated (2S)-2-amino-6-(2,5-dimethylpyrrolidin-1-yl)hexanoic acidUnrated 6-(2,5-dimethylpyrrolidin-1-yl)norleucineUnrated
selenium substitution for sulfurUnrated Se(S)ResUnrated
crosslinked L-aspartic acid residueUnrated XlnkAspUnrated
2-amino-4-(methylthio)butanamideUnrated (2S)-2-amino-4-(methylsulfanyl)butanamideUnrated amidated L-methionineUnrated L-methionine amideUnrated methioninamideUnrated MetNUnrated MOD_RES Methionine amideUnrated
MmtUnrated monomethoxytritylUnrated
(2R)-2-amino-3-([(2S)-2-amino-3-phenylpropanoyl]sulfanyl)propanoic acidUnrated CROSSLNK Phenylalanyl cysteine thioester (Cys-Phe)Unrated phenylalanine cysteine thioesterUnrated S-(2-amino-3-phenylpropanoyl)cysteineUnrated S-(L-phenylalanyl)-L-cysteineUnrated XLNK1PheSCysUnrated
crosslinked L-selenocysteine residueUnrated
1-carboxy-N,N-dimethylaminomethaneUnrated 2-(dimethylamino)acetic acidUnrated (dimethylamino)ethanoic acidUnrated MOD_RES N,N-dimethylglycineUnrated N,N-dimethylglycineUnrated vitamin B16Unrated
2-oxopyrrolidine-5-carboxylic acidUnrated 2-pyrrolidone-5-carboxylic acidUnrated 2-pyrrolidone-5-carboxylic acid (Gln)Unrated (2S)-5-oxo-2-pyrrolidinecarboxylic acidUnrated 5-oxoprolineUnrated 5-oxopyrrolidine-2-carboxylic acidUnrated 5-pyrrolidone-2-carboxylic acidUnrated MOD_RES Pyrrolidone carboxylic acidUnrated N-pyrrolidone carboxyl (N terminus)Unrated ntermpeppyroqUnrated PCAUnrated PyrGlu(Gln)Unrated pyroglutamic acidUnrated Pyroglutamic Acid formed from GlnUnrated PyroglutamylUnrated
MeHisUnrated methylated histidineUnrated
timnodonoylated residueUnrated
2-acetylamino-6-aminohexanoic acidUnrated 2-acetylazanyl-6-azanylhexanoic acidUnrated (2S)-2-acetamido-6-aminohexanoic acidUnrated N2-acetylated L-lysineUnrated N2-acetyl-L-lysineUnrated N2-acetyllysineUnrated N2AcLysUnrated
(2S)-2-amino-6-([(E)-[(3S)-3-amino-3-carboxypropyl](methyl)-lambda(4)-sulfanylidene]amino)hexanoic acidUnrated (E)-N6-([(3S)-3-amino-3-carboxypropyl](methyl)-lambda(4)-sulfanylidene)-L-lysineUnrated S-(L-lysyl)-L-methionine sulfilimineUnrated S-lysyl-methionineUnrated
(2S)-2-amino-6-(dimethylamino)hexanoic acidUnrated dimethylkUnrated epsilon-dimethyllysineUnrated lysine derivative Lys(y)Unrated MOD_RES N6,N6-dimethyllysineUnrated N6Me2LysUnrated N6,N6-dimethylated L-lysineUnrated N6,N6-dimethyl-L-lysineUnrated N(zeta)-dimethyllysineUnrated
2x(13)C,4x(2)H labeled dimethylated L-lysineUnrated
(2S)-2-amino-3-[1-(5-[(2S)-2-amino-2-carboxyethyl]-2-hydroxyphenyl)-1H-imidazol-4-yl]propanoic acidUnrated 3'-(1'-L-histidyl)-L-tyrosineUnrated 3'-(N1'-histidyl)tyrosineUnrated 3'-(N(epsilon)-histidyl)tyrosineUnrated 3'-(tau-histidyl)tyrosineUnrated 3'-(tele-histidyl)tyrosineUnrated CROSSLNK 1'-histidyl-3'-tyrosine (His-Tyr)Unrated
(2S,3R)-2-amino-3-(5-[(2S)-2-amino-2-carboxyethyl]-1H-imidazol-1-yl)-3-(4-hydroxyphenyl)propanoic acidUnrated 3-(3'-L-histidyl)-L-tyrosineUnrated 3-(N3'-histidyl)tyrosineUnrated 3-(pi-histidyl)tyrosineUnrated 3-(pros-histidyl)tyrosineUnrated beta-(N3'-histidyl)tyrosineUnrated beta-(N(delta)-histidyl)tyrosineUnrated CROSSLNK 3'-histidyl-3-tyrosine (His-Tyr)Unrated
N-L-glutamyl-poly-L-glutamic acidUnrated
N6-linoloyl-L-lysineUnrated
Bruker Daltonics SERVA-ICPL(TM) quantification chemistry, medium form - site N-termUnrated
2-(L-cystein-S-yl)-L-phenylalanineUnrated (2R)-2-amino-2-[(2R)-2-amino-2-carboxyethyl]sulfanyl-3-phenylpropanoic acidUnrated (2R,5R)-2,5-diamino-3-thia-2-phenylmethylhexanedioic acidUnrated 2-(S-L-cysteinyl)-L-phenylalanineUnrated alpha-(L-cystein-S-yl)-L-phenylalanineUnrated CROSSLNK 2-cysteinyl-L-phenylalanine (Cys-Phe)Unrated
(2S)-2-amino-3-[4-(5'-adenosine phosphonoxy)phenyl]propanoic acidUnrated 5'-adenylic-O-tyrosineUnrated hydrogen 5'-adenylate tyrosine esterUnrated MOD_RES O-AMP-tyrosineUnrated O4'-L-tyrosine 5'-adenosine phosphodiesterUnrated O4'-(phospho-5'-adenosine)-L-tyrosineUnrated O-5'-Adenosylation ( of Tyrosine)Unrated OAMPTyrUnrated
MeGlnUnrated methylated glutamineUnrated
4-methyl-1-methylidenepiperazin-1-iumUnrated iTRAQ8plex-115 reporter fragmentUnrated
[13-[(S)-(4-amino-4-carboxy)butanoyloxymethyl]-3-[(S)-(3-amino-3-carboxy)propanoyloxymethyl]-7,12-diethenyl-8,17-dimethyl-21H,23H-porphine-2,18-bis(2-carboxyethyl)-N21,N22,N23,N24]-ferrateUnrated 1,5-bishydroxymethyl protoporphyrin IX 1-glutamate ester 5-aspartate esterUnrated BINDING Heme (covalent; via 2 links)Unrated dihydroxyheme-L-aspartate ester-L-glutamate esterUnrated peroxidase heme cofactorUnrated
9x(13)C labeled residueUnrated
ethylated residueUnrated EtResUnrated
nickel containing modified residueUnrated NiResUnrated
(2S)-2-amino-3-hydroxy-3-(1H-imidazol-4-yl)propanoic acidUnrated 3-hydroxy-L-histidineUnrated MOD_RES (3S)-3-hydroxyhistidineUnrated
MeLysUnrated methylated lysineUnrated
(2S,3R)-2-amino-3-(riboflavin 5'-hydrogen phosphonoxy)butanoic acidUnrated MOD_RES FMN phosphoryl threonineUnrated O3-threonyl flavin mononucleotideUnrated O3-threonyl FMNUnrated OFMNThrUnrated O-(riboflavin phosphoryl)-L-threonineUnrated
deamidated 4-methyl esterified asparagineUnrated
L-selenocysteinyl molybdenum bis(molybdopterin guanine dinucleotide) (Sec)Unrated
2-[1-azanyl-2-phenylethyl]-4,5-dihydro-1,3-thiazole-4-carboxylic acidUnrated (4R)-2-[(1S)-1-amino-2-phenylethyl]-4,5-dihydro-1,3-thiazole-4-carboxylic acidUnrated CROSSLNK Thiazoline-4-carboxylic acid (Phe-Cys)Unrated L-phenylalanine thiazoline-4-carboxylic acidUnrated
2-(acetylamino)butanedioic acidUnrated 2-(acetylazanyl)butanedioic acidUnrated (2S)-2-(acetamido)butanedioic acidUnrated AcAspUnrated acetylaspartic acidUnrated MOD_RES N-acetylaspartateUnrated N-acetylated L-aspartic acidUnrated N-acetyl-L-aspartic acidUnrated
4x(2)H labeled dimethylated L-lysineUnrated mod189Unrated
nitrated L-tyrosineUnrated nitroyUnrated
crosslinked L-glutamine residueUnrated XlnkGlnUnrated
15,16-DhbvUnrated 15,16-dihydrobiliverdin cysteine adductUnrated 15,16-dihydrobiliverdin IXalphaUnrated (16R)-18-ethenyl-8,12-bis(2-carboxyethyl)-3-[(1R)-1-(((2R)-2-amino-2-carboxy)ethylsulfanyl)ethyl]-2,7,13,17-tetramethyl-15,16-dihydrobilin-1,19(21H,24H)-dioneUnrated 18-ethenyl-8,12-bis(2-carboxyethyl)-3-(2-(cysteinyl-S)-ethyl)-2,7,13,17-tetramethylbiladiene-ab-1,19(16H,21H)-dioneUnrated 3alpha-cysteinyl-15,16-dihydrobiliverdinUnrated 3'-cysteinyl-15,16-dihydrobiliverdinUnrated BINDING 15,16-dihydrobiliverdin (covalent; via 1 link)Unrated DBVUnrated S-15,16-dihydrobiliverdin-L-cysteineUnrated
biotin polyethyleneoxide amineUnrated
Me1HisUnrated methylhUnrated MOD_RES MethylhistidineUnrated monomethylated L-histidineUnrated
2-amino-5-guanidinohexanoic acidUnrated (2S,5S)-2-amino-5-carbamimidamidohexanoic acidUnrated 5-methylated L-arginineUnrated 5-methyl-L-arginineUnrated C5Me1ArgUnrated delta-methylarginineUnrated MOD_RES 5-methylarginineUnrated
2-amino-4-(methylthio)butanoic acidUnrated 2-amino-4-(methylthio)butyric acidUnrated (2R)-2-amino-4-(methylsulfanyl)butanoic acidUnrated D-MetUnrated D-methionineUnrated MOD_RES D-methionineUnrated
(2S)-2-amino-6-[(2R)-2-hydroxy-3-(phosphonooxy)propanamido]hexanoic acidUnrated (2S)-2-amino-6-([(2R)-2-hydroxy-3-(phosphonooxy)propanoyl]amino)hexanoic acidUnrated 3-phosphoglyceryl-lysineUnrated N6-(3-phosphoglyceryl)-L-lysineUnrated
2-hydroxypropionic acidUnrated (2R)-2-hydroxypropanoic acidUnrated alpha-hydroxypropionic acidUnrated Lac(Ser)Unrated lactic acidUnrated MOD_RES Lactic acidUnrated
mono N-acetylated residueUnrated N-AcetylUnrated N-AcetylationUnrated NAcResUnrated
2-[(1S)-1-amino-2-hydroxyethyl]-1,3-oxazole-4-carboxylic acidUnrated 2-[(1S)-1-azanyl-2-hydroxyethyl]-1,3-oxazole-4-carboxylic acidUnrated CROSSLNK Oxazole-4-carboxylic acid (Ser-Ser)Unrated L-serine oxazole-4-carboxylic acidUnrated
(2S,4S)-2,4-diamino-2-[([(2R)-2-amino-2-carboxyethyl]sulfanyl)methyl]pentanedioic acidUnrated (2S,4S,8R)-2,4,8-triamino-4-carboxy-6-thianonanedioic acidUnrated (2S,4S,8R)-labioninUnrated labioninUnrated
(18)O monosubstituted L-tyrosineUnrated
2-azanyl-3-hydroxybutanoic acidUnrated (2S,3R)-2-amino-3-hydroxybutanoic acidUnrated alpha-amino-beta-hydroxybutyric acidUnrated beta-methylserineUnrated L-threonineUnrated L-threonine residueUnrated L-threo-threonineUnrated ThrUnrated
succinylated residueUnrated
3-[(2,6-dimethylpiperidin-1-yl)acetyl]amino)propanoylUnrated TMT6plex-131 reporter+balance reagent acylated N-terminalUnrated
N6-ethyl-L-lysineUnrated
OPhUnrated phenyl esterUnrated
APTAUnrated

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