Protein Modification Ontology (MOD)

Name: Protein Modification Ontology (MOD)
Published: 2026-11-13T13:50:45.374984Z

Selected English labels and exact synonyms from the Protein Modification Ontology (MOD), retrieved 2026-11-07, with source identifiers and spelling preserved. Licensed CC BY 4.0; full attribution is listed under Sources. Under semantic review; not endorsed by the ontology's maintainers.

Curator: Oliver Bennett

Languages shown reflect this page; source translations may cover only some concepts.

English
GSIAsnUnrated LIPID GPI-like-anchor amidated asparagineUnrated N-asparaginyl-glycosylsphingolipidinositolethanolamineUnrated
2-azanyl-3-(4-hydoxyphenyl)propanoic acidUnrated (2S)-2-amino-3-(4-hydoxyphenyl)propanoic acidUnrated alpha-amino-beta-(para-hydroxyphenyl)propionic acidUnrated L-tyrosineUnrated L-tyrosine residueUnrated para-hydroxyphenylalanineUnrated p-tyrosineUnrated TyrUnrated
O3-(ADP-ribosyl)-L-threonineUnrated O3-[alpha-D-ribofuranoside 5'->5'-ester with adenosine 5'-(trihydrogen diphosphate)]-L-threonineUnrated O3-alpha-D-ribofuranosyl-L-threonine 5'->5'-ester with adenosine 5'-(trihydrogen diphosphate)Unrated O-(ADP-ribosyl)-L-threonineUnrated (S)-2-amino-3-([adenosine 5'-(trihydrogen diphosphate) 5'->5'-ester with alpha-D-ribofuranosyl]oxy)-butanoic acidUnrated
flavin crosslinked residuesUnrated
piperidine adduct to C-terminal CysUnrated
Me2+ResUnrated protonated-dimethylated residueUnrated
(2-aminoacetyl)oxy-([(2R,3R,4R,5R)-5-(6-aminopurin-9-yl)-3,4-dihydroxy-oxolan-2-yl]methoxy)phosphinic acidUnrated ([(2R,3S,4R,5R)-5-(6-aminopurin-9-yl)-3,4-dihydroxy-oxolan-2-yl]methoxy-hydroxy-phosphoryl)-2-aminoethanoateUnrated 5'-adenylic-glyinateUnrated aminoacetyl [5-(6-amino-9H-purin-9-yl)-3,4-dihydroxytetrahydrofuran-2-yl]methyl hydrogen phosphateUnrated glycine monoanhydride with 5'-adenylic acidUnrated glycyl 5'-adenylateUnrated glycyl adenosine-5'-phosphateUnrated glycyladenylateUnrated glycyl phospho-5'-adenosineUnrated MOD_RES Glycyl adenylateUnrated
2-([4-azanyl-4-carboxybutanoyl]azanyl)pentanedioic acidUnrated (2S)-2-[(4S)-4-amino-4-carboxybutanamido]pentanedioic acidUnrated (2S)-2-([(4S)-4-amino-4-carboxybutanoyl]amino)pentanedioic acidUnrated 5-glutamyl glutamic acidUnrated gamma-glutamylglutamateUnrated isoglutamyl glutamic acidUnrated isoglutamyl monoglutamic acidUnrated MOD_RES 5-glutamyl glutamateUnrated N alpha -(gamma-Glutamyl)-GluUnrated N-(gamma-L-glutamyl)-L-glutamic acidUnrated
HexNAc2 N4-glycosylated asparagineUnrated
5GlyceroPEtAGluUnrated 5-L-glutamyl glycerylphosphorylethanolamineUnrated L-glutamyl 5-glycerophosphoethanolamineUnrated L-glutamyl 5-glycerophosphorylethanolamineUnrated L-glutamyl 5-glycerylphosphorylethanolamineUnrated MOD_RES 5-glutamyl glycerylphosphorylethanolamineUnrated (S)-2-amino-5-[2-([([2,3-dihydroxypropyl]oxy)(hydroxy)phosphoryl]oxy)ethyl]amino-5-oxopentanoic acidUnrated
MeThrUnrated methylated threonineUnrated
MOD_RES FMN phosphoryl serineUnrated O3-seryl flavin mononucleotideUnrated O3-seryl FMNUnrated OFMNSerUnrated O-(riboflavin phosphoryl)-L-serineUnrated (R)-2-amino-3-(riboflavin 5'-hydrogen phosphonoxy)propanoic acidUnrated
2-[1,5-bis(azanyl)pentyl]-1,3-thiazole-4-carboxylic acidUnrated 2-[(1S)-1,5-diaminopentyl]-1,3-thiazole-4-carboxylic acidUnrated L-lysine thiazole-4-carboxylic acidUnrated
hydroxylated lysineUnrated HyLysUnrated
(12-ethenyl-7-[(1S)-1-([(2R)-2-amino-2-carboxyethyl]sulfanyl)ethyl]-3,8,13,17-tetramethyl-21H,23H-porphine-2,18-bis[2-carboxyethyl]-N21,N22,N23,N24)-ferrateUnrated 4-[1-(S-cysteinyl)ethyl]protoporphyrin IXUnrated BINDING Heme (covalent; via 1 link)Unrated cysteinyl hemeUnrated HemeCys1Unrated heme-L-cysteineUnrated S-Heme (on Cysteine)Unrated
5-hydroxy-L-tryptophanUnrated (S)-2-Amino-3-(5-hydroxy-1H-indol-3-yl)propanoic acidUnrated
2-amino-6-propionylaminocaproic acidUnrated (2S)-2-amino-6-(propanoylamino)hexanoic acidUnrated epsilon-propanoyl-L-lysineUnrated epsilon-propionyl-L-lysineUnrated MOD_RES N6-propionyllysineUnrated N6-(1-oxopropyl)-L-lysineUnrated N6-propanoyl-L-lysineUnrated N6-propionyllysineUnrated N(zeta)-propanoyllysineUnrated
(15)N labeled residueUnrated
(2S)-2-amino-3-(5-chloro-1H-indol-3-yl)propanoic acidUnrated 5'-chloro-L-tryptophanUnrated MOD_RES 5'-chlorotryptophanUnrated
L-methionine removalUnrated ntermmcleaveUnrated
(2R,3R)-2-amino-3-hydroxybutanoic acidUnrated D-allo-threonineUnrated D-ThrUnrated D-threonineUnrated MOD_RES D-threonineUnrated
reduced lysine 4-hydroxynonenal adductUnrated
Levuglandinyl - hydroxylactam adduct, K and N-termUnrated
thiophosphate labeled with biotin-HPDPUnrated
Hy3ResUnrated trihydroxylated residueUnrated
(2R)-2-amino-3-([2-([(methylsulfanyl)carbonyl]amino)ethyl]sulfanyl)propanoic acidUnrated L-cysteine N-(methylthiocarbonyl)aziridine adductUnrated MTCTKUnrated N6-methylthiocarbonyl-4-thialysineUnrated S-[2-([(methylthio)carbonyl]amino)ethyl]-L-cysteineUnrated S-methylthiocarbonylaminoethylcysteine (Cys)Unrated
Me3ResUnrated trimethylated residueUnrated
(2S)-3-hydroxy-2-(methylamino)propanoic acidUnrated MOD_RES N-methylserineUnrated N-methyl-L-serineUnrated N-methylserineUnrated
monohydroxylated phenylalanineUnrated
pyridylacetylated residueUnrated
3-(2-methylsulfanyl)ethyl-6-(4-hydroxybenzylidene)-5-iminopiperazin-2-oneUnrated 3-(2-methylthio)ethyl-6-(4-hydroxybenzylidene)-5-amino-3,6-didehydropyrazin-2-olUnrated 3-(2-methylthio)ethyl-6-(4-hydroxybenzylidene)-5-iminopiperazin-2-oneUnrated GFP-like chromoprotein asFP595 chromophoreUnrated L-methionyl-L-tyrosyl-2-keto-5-iminopiperazineUnrated
2-amino-6-butyrylaminocaproic acidUnrated (2S)-2-amino-6-(butanoylamino)hexanoic acidUnrated (2S)-2-azanyl-6-(butanoylamino)hexanoic acidUnrated epsilon-butanoyl-L-lysineUnrated epsilon-butyryl-L-lysineUnrated MOD_RES N6-butyryllysineUnrated N6-(1-oxobutyl)-L-lysineUnrated N6-butanoyl-L-lysineUnrated N6-butyryllysineUnrated N(zeta)-butanoyllysineUnrated
glycoconjugated residueUnrated
MeMetUnrated methylated methionineUnrated
(2S,3R)-2-amino-3-(hexadecanoyloxy)butanoic acidUnrated LIPID O-palmitoyl threonineUnrated L-threonine hexadecanoate esterUnrated O3-palmitoyl-threonineUnrated O-hexadecanoylated L-threonineUnrated O-palmitoylated L-threonineUnrated O-palmitoyl-L-threonineUnrated OPamThrUnrated palmitoylationtUnrated threonine palmitate esterUnrated
1-azanyl-4-(4-[2-azanyl-2-carboxyethyl]-1H-imidazol-2-yl)-N,N,N-trimethyl-1-oxobutan-2-azaniumUnrated 2-[3-carboxamido-3-(trimethylammonio)propyl]histidineUnrated 2'-[3-carboxamido-3-(trimethylammonio)propyl]-L-histidineUnrated 2-amino-3-[[2-(3-amino-3-carbamoyl-prop-1-enyl)-1,1,3-trimethyl-2,3-dihydroimidazol-5-yl]]propanoic acidUnrated 2-amino-4-[[5-(2-amino-2-carboxylato-ethyl)-1,1,3-trimethyl-2,3-dihydroimidazol-2-yl]]but-3-enamideUnrated (2R)-1-amino-4-(4-[(2S)-2-amino-2-carboxyethyl]-1H-imidazol-2-yl)-N,N,N-trimethyl-1-oxobutan-2-aminiumUnrated 2-[(R)-3-carboxamido-3-(trimethylammonio)propyl]-4-((S)-2-amino-2-carboxyethyl)-1H-imidazoleUnrated (3-[4-(2-amino-2-carboxy-ethyl)-1H-imidazol-2-yl]-1-carbamoyl-propyl)-trimethylammoniumUnrated alpha-(aminocarbonyl)-4-(2-amino-2-carboxyethyl)-N,N,N-trimethyl-1H-imidazole-2-propanaminiumUnrated DiphthUnrated diphthamideUnrated diphthamide (from histidine)Unrated MOD_RES DiphthamideUnrated
Double oxidation of TrpUnrated formylkynureninwUnrated N'-formyl-L-kynurenineUnrated
GalNAcResUnrated mono-N-acetylaminogalactosylated residueUnrated
metal or metal cluster containing modified residueUnrated MetalResUnrated
alpha-carboxyl deprotonated residueUnrated C1H-NResUnrated
vanadium containing modified residueUnrated VResUnrated
Hex1HexNAc1NeuAc1 O-glycosylated threonineUnrated
3-[(2,6-dimethylpiperidin-1-yl)acetyl]amino)propanoylUnrated TMT6plex-129 reporter+balance reagent O3-acylated serineUnrated
3-[(2,6-dimethylpiperidin-1-yl)acetyl]amino)propanoylUnrated TMT6plex-126 reporter+balance reagent N'-acylated histidineUnrated
sulfur containing modified residueUnrated
(4-methylpiperazin-1-yl)acetylUnrated iTRAQ8plex-118 reporter+balance reagent O3-acylated threonineUnrated
2-aminosuccinimide ring crosslinked residuesUnrated
crosslinked L-alanine residueUnrated XlnkAlaUnrated
crosslinked residues with loss of ammoniaUnrated
2-(hexadecanamido)-4-(methylsulfanyl)butanoic acidUnrated (2S)-2-(hexadecanoylamino)-4-(methylsulfanyl)butanoic acidUnrated LIPID N-palmitoyl methionineUnrated N-(1-oxohexadecyl)methionineUnrated N-palmitoyl-L-methionineUnrated
SGlycoResUnrated S-glycosylated residueUnrated
L-tryptophan removalUnrated
pyridyl thiol modified L-threonineUnrated
L-isoglutamyl triglutamic acidUnrated N alpha -(gamma-Glutamyl)-Glu3Unrated
2dHdH2OResUnrated didehydrogenated and dehydrated residueUnrated
dihydroxylated isoleucineUnrated Hy2IleUnrated
alpha-amino pyridylacetylated residueUnrated ntermpeppyridylUnrated
chondroitin 4-sulfate (chondroitin sulfate A)Unrated chondroitin 6-sulfate (chondroitin sulfate C)Unrated chondroitin sulfate D-glucuronosyl-D-galactosyl-D-galactosyl-D-xylosyl-L-serineUnrated poly[beta-1,4-D-glucopyranuronosyl-beta-1,3-(2-acetamido-2-deoxy-4-sulfate-D-galactosyl)]-beta-1,4-D-glucopyranuronosyl-beta-1,3-D-galactosyl-beta-1,3-D-galactosyl-beta-1,4-D-xylosyl-beta-1,3-L-serine; poly[beta-1,4-D-glucopyranuronosyl-beta-1,3-(2-acetamido-2-deoxy-6-sulfate D-galactosyl)]-beta-1,4-D-glucopyranuronosyl-beta-1,3-D-galactosyl-beta-1,3-D-galactosyl-beta-1,4-D-xylosyl-beta-1,3-L-serineUnrated
2-amino-3-(2-hydroxyphenyl)propanoic acidUnrated 2-hydroxy-L-phenylalanineUnrated DL-o-TyrosineUnrated o-tyrosineUnrated O-TyrosineUnrated
cysteinyl mercuryUnrated
oxidized proline biotinylated with biotin-LC-hydrazide, reducedUnrated
formyl tryptophanylUnrated N-formyl-L-tryptophanUnrated
gamma-aminobutyrylUnrated
(4-methylpiperazin-1-yl)acetylUnrated iTRAQ8plex reporter+balance reagent acylated N-terminalUnrated
O-hexadecanoylated residueUnrated O-palmitoylated residueUnrated OPamResUnrated
beta-heparinUnrated chondroitin sulfate BUnrated dermatan 4-sulfate D-glucuronosyl-D-galactosyl-D-galactosyl-D-xylosyl-L-serineUnrated poly[beta-1,4-L-idopyranuronosyl-alpha-1,3-(2-acetamido-2-deoxy-4-sulfate D-galactosyl)]-beta-1,4-D-glucopyranuronosyl-beta-1,3-D-galactosyl-beta-1,3-D-galactosyl-beta-1,4-D-xylosyl-beta-1,3-L-serineUnrated
2-formamidopentanedioic acidUnrated 2-formylaminopentanedioic acidUnrated 2-(formylazanyl)pentanedioic acidUnrated (2S)-2-(formylamino)pentanedioic acidUnrated N-formyl-L-glutamic acidUnrated
Hex2Unrated
(4-methylpiperazin-1-yl)acetylUnrated iTRAQ8plex-115 reporter+balance reagent O4'-acylated tyrosineUnrated
Me1AspUnrated methyldUnrated monomethylated L-aspartic acidUnrated
2-amino-3-(4-hydroxyphenyl)propanoic acid 4'-sulfateUnrated MOD_RES SulfotyrosineUnrated O4'-sulfo-L-tyrosineUnrated O4-sulfotyrosineUnrated OSulfTyrUnrated (S)-2-amino-3-(4-sulfooxyphenyl)propanoic acidUnrated sulfationyUnrated Sulphation (of O of Tyrosine)Unrated tyrosine-O-sulfonic acidUnrated tyrosine-O-sulphonic acidUnrated tyrosine sulfateUnrated Tyrosinyl SulphateUnrated
4-methyl-1-methylidenepiperazin-1-iumUnrated iTRAQ8plex-117 reporter fragmentUnrated
2-[([(2S)-2-amino-2-carboxyethoxy][hydroxy]phosphoryl)oxy]-N,N,N-trimethylethanaminiumUnrated 2-[([(2S)-2-azanyl-2-carboxyethoxy][hydroxy]phosphoryl)oxy]-N,N,N-trimethylethanazaniumUnrated MOD_RES O-(2-cholinephosphoryl)serineUnrated O3-[(2-[trimethylammonio]ethyl)phosphoryl]-L-serineUnrated O3-phosphocholine-L-serineUnrated O-cholinephosphoryl-L-serineUnrated serine choline phosphateUnrated serine choline phosphodiesterUnrated
2,3-trans-3,4-trans-3,4-dihydroxy-L-prolineUnrated 2-alpha-3-beta-4-alpha-3,4-dihydroxyprolineUnrated (2S,3R,4R)-3,4-dihydroxyprolineUnrated (2S,3R,4R)-3,4-dihydroxypyrrolidine-2-carboxylic acidUnrated MOD_RES (3R,4R)-3,4-dihydroxyprolineUnrated
cysteine monobromobimane derivativeUnrated
3-[(2,6-dimethylpiperidin-1-yl)acetyl]amino)propanoylUnrated TMT6plex-127 reporter+balance reagent acylated N-terminalUnrated
(2S)-2-amino-3-([(Z)-2-aminoethenyl]sulfanyl)propanoic acidUnrated CROSSLNK S-(2-aminovinyl)-D-cysteine (Ser-Cys)Unrated S-(2-aminovinyl)-D-cysteineUnrated S-(2-aminovinyl)-D-cysteine (Cys-Ser)Unrated (S,Z)-S-(2-aminovinyl)cysteineUnrated XLNK-(D)SCys-VinAmUnrated
GPIAspUnrated LIPID GPI-anchor amidated aspartateUnrated N-aspartyl-glycosylphosphatidylinositolethanolamineUnrated
3-[(2,6-dimethylpiperidin-1-yl)acetyl]amino)propanoylUnrated TMT6plex-131 reporter+balance reagent O3-acylated serineUnrated
2-amino-3-([2-aminoethenyl]sulfanyl)propanoic acidUnrated S-(2-aminovinyl)-cysteineUnrated
prompt loss of methanethiol from oxidixed methionineUnrated
GlcUnrated glucosylated residueUnrated
2-azanyl-6-[(3-carboxypropanoyl)azanyl]hexanoic acidUnrated (2S)-2-amino-6-[(3-carboxypropanoyl)amino]hexanoic acidUnrated 4-[[(5S)-5-amino-6-hydroxy-6-oxohexyl]amino]-4-oxobutanoateUnrated MOD_RES N6-succinyllysineUnrated N6-succinyl-L-lysineUnrated N(epsilon)-(succinyl)lysineUnrated succinyllysineUnrated
1x(13)C,3x(2)H labeled monomethylated residueUnrated
2-azanyl-3-methylpentanoic acidUnrated (2S,3S)-2-amino-3-methylpentanoic acidUnrated 3-methyl-norvalineUnrated alpha-amino-beta-methylvaleric acidUnrated IleUnrated IsoleucylUnrated L-erythro-isoleucineUnrated L-isoleucineUnrated L-isoleucine residueUnrated
EDT-iodo-PEO-biotinUnrated
(2S)-2-[(3R,4R)-3,4-dihydroxy-5-(phosphonooxy)pentylidene]amino-3-methylbutanoic acidUnrated ACT_SITE Schiff-base intermediate with DNA; via amino nitrogenUnrated DNA glycosylase valine Schiff base intermediateUnrated N-(DNA-1',2'-dideoxyribos-1'-ylidene)-L-valineUnrated
HexAUnrated hexosuronylated residueUnrated
(2S)-2-amino-4-(dimethylsulfonio)butanoateUnrated (3S)-(3-amino-3-carboxypropyl)dimethylsulfaniumUnrated [(3S)-3-amino-3-carboxypropyl](dimethyl)sulfoniumUnrated S-methyl-L-methionineUnrated S-methylmethionineUnrated S-methylmethioniniumUnrated vitamin UUnrated
cervonoylated residueUnrated
O-thiophospho-L-threonineUnrated
2-amino-4-methylpentanoic methyl esterUnrated alpha-aminoisocaproic methyl esterUnrated L-leucine methyl esterUnrated methyl (2S)-2-amino-4-methylpentanoateUnrated methyl esterified L-leucineUnrated methyl L-leucinateUnrated MOD_RES Leucine methyl esterUnrated OMeLeuUnrated
modified L-selenocysteine residueUnrated
2-[(1S)-1-amino-4-([diaminomethylidene]amino)butyl]-1,3-thiazole-4-carboxylic acidUnrated CROSSLNK Thiazole-4-carboxylic acid (Arg-Cys)Unrated L-arginine thiazole-4-carboxylic acidUnrated
(4-methylpiperazin-1-yl)acetylUnrated iTRAQ4plex-114 reporter+balance reagent acylated residueUnrated
propanoylated residueUnrated
2,6-diamino-3-methyl-4-thiaheptanedioic acidUnrated 2-amino-3-([2-amino-2-carboxyethyl]sulfanyl)butanoic acidUnrated 2-amino-3-[(2-amino-2-carboxyethyl)thio]butanoic acidUnrated 2-azanyl-3-[(2-azanyl-2-carboxyethyl)sulfanyl]butanoic acidUnrated 3-methyllanthionineUnrated cysteine-3-L-butyrine thioetherUnrated
Hex1HexNAc2dHex1 N4-glycosylated asparagineUnrated
(2S)-2-([(4S)-4-amino-4-carboxybutanoyl]amino)hexanedioic acidUnrated 5-glutamyl 2-aminoadipic acidUnrated MOD_RES 5-glutamyl 2-aminoadipic acidUnrated N2-(gamma-glutamyl)-2-aminoadipic acidUnrated N2-(isoglutamyl)-2-aminoadipic acidUnrated
cholesteryl glycinateUnrated glycine cholest-5-en-3beta-ol esterUnrated glycine cholesterol esterUnrated hedgehog lipophilic adductUnrated LIPID Cholesterol glycine esterUnrated

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